I do not believe so. They way it was explained is that the unit will only benefit from the blessing's stat boost. So, during Aether Raids Eir will grant her + Res and HP boost to Legendary Heroes no matter their element so long as Light Season is in play.
Then the second part, is that Mythic Heroes like Eir will receive the stat boosts from any Legendary Hero whose element matches one in the current season, so during Fire Season Eir would receive a boost to HP and Atk if paired with Legendary Hector in Arena.
Now, in Arena Assault, I think any unit with a blessing or Legendary unit gets those extra points for kills, but in regular Arena only the bonus units will get the extra points for kills.
Anyway, that is how I understood the explanation, but I could be incorrect. I'm not 100% certain if both elements have to be in the current season to benefit or if they have to be "matched pairs" i.e. Light & Water only; Wind & Darkness only; Earth & Aether only; and Fire & Animus only.
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